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[Updated 2/1] The configuration shown here is improved, and appears to have solved the problem. I have upgraded the Fume-shroom that was in rows 2 and 5, column 6 to a Gloom-shroom. The extra "backdraft" from that seems to be keeping the Pumpkins around the Cattails healthy.
However, I still don't what was eating them, as I have yet to see any zombies make it that far down the pool.